1986 - PhD in biophysics, University of California, Berkeley, USA
1987 - Post doctoral fellow University of California, San Francisco, USA
1993 - Group Leader, MRC-LMB, Cambridge, UK
1997 - Group leader, MRC NIMR, London, UK
2006 - Elected member of EMBO
2010 - Elected fellow of the Academy of Medical Sciences
2012 - Joint head of division, NIMR, London, UK
2013 - Elected fellow of the Royal Society
2015 - Group Leader, the Francis Crick Institute, London
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